# Beef Cattle Profitability: Factors, Benchmarks, and Improvement Strategies


## Key Takeaways

- Beef cattle profitability is fundamentally driven by the relationship between cost of production per head and revenue per head, with feed costs representing 60-70% of total production expenses.
- Optimal reproductive performance, measured by a calving rate of 85-95% of exposed females, is critical for spreading fixed costs over a larger number of calves sold.
- Weaning weights within the 500-650 lbs (227-295 kg) benchmark at 205 days, influenced by genetics, nutrition, and health, directly increase revenue per head when market prices are favorable.
- Feed efficiency, quantified by a feed conversion ratio of 6.0-7.5 lbs feed per lb gain, is paramount in feedlot operations to reduce the largest single expense and improve the cost of gain.
- Strategic market timing, considering seasonal price patterns and optimal sale weights (550-750 lbs for feeder calves, 1,200-1,400 lbs for finished cattle), is essential for maximizing revenue per pound.
- Comprehensive record-keeping and benchmarking against industry standards, such as the Standardized Performance Analysis (SPA) program, are vital for identifying specific areas for improvement and making informed management decisions.

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Beef cattle profitability depends on the relationship between production costs per head and revenue per head, which is determined by weaning weight, feed efficiency, market timing, and reproductive performance. Producers who track these factors against established benchmarks can identify specific areas for improvement and make informed management decisions. This article provides benchmark ranges for cost of production per head, outlines the key profitability drivers, and offers practical strategies for improving financial performance in beef cattle operations.

## At a Glance: Key Profitability Drivers and Benchmarks

The table below summarizes the primary factors influencing beef cattle profitability and provides benchmark ranges that producers can use to evaluate their own operation's performance.

| Profitability Driver | Benchmark Range | Impact on Profitability |
|----------------------|-----------------|------------------------|
| Weaning weight per calf | 500-650 lbs (227-295 kg) at 205 days | Higher weaning weights increase revenue per head when market prices are favorable |
| [Feed conversion ratio](/knowledge/animal-farming/poultry/feed-conversion-ratio-measuring-improving-poultry-efficiency) (feedlot phase) | 6.0-7.5 lbs feed per lb gain | Lower ratios reduce feed costs, which represent 60-70% of total production costs |
| Calving rate (pregnancy rate) | 85-95% of exposed females | Higher calving rates spread fixed costs over more calves sold |
| Cost of production per head | Varies by region and system, typical range $800-$1,200 per cow per year | Directly determines breakeven price and margin per head |
| Market timing (sale weight and season) | 550-750 lbs for feeder calves, 1,200-1,400 lbs for finished cattle | Timing affects price per pound and total revenue |

## Understanding the Cost of Production per Head

Cost of production per head is the single most important metric for assessing beef cattle profitability. It includes all expenses incurred to maintain a breeding female and produce a weaned calf, or to feed an animal through the stocker or feedlot phase. Without accurate cost tracking, producers cannot determine whether their operation is generating a profit or a loss.

### Fixed Costs

Fixed costs do not change with the number of animals and must be covered regardless of production levels. These include land payments or lease costs, facility depreciation, equipment maintenance, and insurance. In many operations, fixed costs account for 20-30% of total annual expenses per cow.

### Variable Costs

Variable costs change directly with herd size and production intensity. Major variable costs include feed and forage, veterinary services and pharmaceuticals, mineral supplements, breeding fees or bull purchases, fuel and utilities, and labor. Feed costs typically represent the largest single expense, often 50-70% of total variable costs depending on the production system.

### Calculating Breakeven Price

The breakeven price is the market price per pound needed to cover all production costs. It is calculated by dividing total annual cost per cow by the pounds of calf sold per cow. For example, if annual cost per cow is $1,000 and the operation sells 500 pounds of calf per cow, the breakeven price is $2.00 per pound. Producers should calculate their breakeven price before making marketing decisions and compare it to current and projected market prices.

## Weaning Weight and Calf Performance

Weaning weight directly affects revenue per head and is influenced by genetics, nutrition, and management. The USDA Natural Resources Conservation Service provides technical guidance on forage management and grazing systems that support optimal calf growth (www.nrcs.usda.gov). Producers should aim for weaning weights within the benchmark range of 500-650 pounds at 205 days of age, adjusted for breed and environment.

### Genetic Selection for Growth

Selecting bulls with high expected progeny differences (EPDs) for weaning weight and yearling weight can improve calf growth rates. However, producers must balance growth traits with maternal traits such as calving ease and milk production. The Merck Veterinary Manual offers guidance on breeding soundness evaluations and genetic selection for beef cattle (www.merckvetmanual.com/management-and-nutrition). Overemphasis on growth without considering feed resources and cow maintenance requirements can reduce overall profitability.

### Nutrition and Forage Management

Calf weaning weight is heavily dependent on the dam's milk production and the quality of forage available to the calf. Adequate protein and energy in the cow's diet during late gestation and early lactation support higher milk yields and calf growth. Producers should test forage quality annually and supplement as needed to meet nutritional requirements. The Food and Agriculture Organization of the United Nations provides resources on sustainable livestock production systems that integrate forage management with animal nutrition (www.fao.org/animal-production/en).

### Health Management

Calf health directly affects weaning weight and subsequent performance. Parasite burdens, respiratory disease, and nutritional deficiencies can reduce growth rates by 10-30%. A 2023 study published in Animals titled "The Economic Impact of Parasitism from Nematodes, Trematodes and Ticks on [Beef Cattle Production](/knowledge/animal-farming/beef-cattle/beef-cattle-production-systems-economics-and-sustainability)" documents the significant economic losses caused by internal and external parasites in beef cattle operations (https://pubmed.ncbi.nlm.nih.gov/37238028). Producers should implement strategic deworming programs based on fecal egg counts and local parasite prevalence data.

## Feed Efficiency and Cost Management

Feed efficiency is the ratio of feed consumed to weight gained. Improving feed efficiency reduces the cost of gain and increases profitability, particularly in feedlot operations where feed costs are the largest expense.

### Measuring Feed Efficiency

[Feed conversion ratio](/knowledge/animal-farming/poultry/feed-conversion-ratio-measuring-improving-poultry-efficiency) (FCR) is calculated as pounds of feed consumed per pound of gain. An FCR of 6.0 means the animal consumed 6 pounds of feed for each pound of weight gained. Residual feed intake (RFI) is a more precise measure that accounts for maintenance requirements and is moderately heritable. Producers can select for improved feed efficiency using genomic testing and performance records.

### Strategies to Improve Feed Efficiency

- Use high-quality forages and balanced rations that meet but do not exceed nutrient requirements
- Group animals by size and condition to reduce competition and allow targeted feeding
- Minimize feed waste through proper feeder design and management
- Select cattle with genetic potential for efficient feed conversion
- Monitor body condition scores and adjust rations accordingly

### Feed Cost Reduction

Producers can reduce feed costs by maximizing the use of homegrown forages, improving pasture utilization through rotational grazing, and purchasing feed ingredients in bulk when prices are favorable. The USDA National Agricultural Library provides information on animal health and welfare practices that support efficient feed utilization (www.nal.usda.gov/animal-health-and-welfare). Overfeeding concentrates or supplements increases costs without proportional gains in performance.

## Market Timing and Price Risk Management

Market timing involves deciding when to sell cattle to capture the best prices. Feeder cattle prices vary seasonally and cyclically, and producers who understand these patterns can improve profitability.

### Seasonal Price Patterns

Feeder cattle prices typically peak in spring and early summer when grass is abundant and demand for stocker cattle is high. Prices often decline in fall as more calves come to market. Finished cattle prices follow different patterns influenced by beef demand, feed costs, and processing capacity. Producers should study historical price data for their region and market class.

### Price Risk Management Tools

- Forward contracts with feedlots or packers
- Futures and options contracts on the Chicago Mercantile Exchange
- Livestock Risk Protection (LRP) insurance subsidized by the USDA
- Cooperative marketing groups that aggregate cattle for better pricing

### Optimal Sale Weight

The optimal sale weight depends on the production system and market conditions. Feeder calves sold at 550-650 pounds often command higher prices per pound than heavier calves, but total revenue per head is lower. Feedlot operators must balance the cost of gain against expected slaughter prices to determine the optimal finish weight. Producers should calculate their breakeven price at different sale weights and compare to projected market prices.

## Reproductive Performance and Herd Productivity

Reproductive performance is the foundation of beef cattle profitability. A cow that does not wean a calf each year generates no revenue but incurs full annual costs. The calving rate, defined as the percentage of exposed females that wean a calf, directly affects the number of calves available for sale and the cost per calf produced.

### Key Reproductive Metrics

- Pregnancy rate: percentage of females exposed to a bull that become pregnant
- Calving rate: percentage of females exposed that wean a live calf
- Calving interval: days between successive calvings (target 365 days)
- Age at first calving: target 22-24 months for heifers

### Improving Reproductive Performance

A 2016 study published in Theriogenology titled "Fertility management of bulls to improve beef cattle productivity" emphasizes the importance of bull fertility in achieving high pregnancy rates (https://pubmed.ncbi.nlm.nih.gov/27173954). Producers should conduct breeding soundness examinations on all bulls 30-60 days before the breeding season. Heifers should be developed to reach 65% of mature body weight before breeding to ensure adequate pelvic size and reproductive maturity.

### Genetics and Genomics

A 2014 study published in Animal titled "Genetics and genomics of reproductive performance in dairy and beef cattle" reviews the genetic basis of fertility traits and the potential for genomic selection to improve reproductive efficiency (https://pubmed.ncbi.nlm.nih.gov/24703258). Genomic testing can identify heifers and bulls with superior genetic potential for fertility, reducing the need for prolonged breeding seasons and culling.

### Culling Decisions

Cows that fail to wean a calf in two consecutive years should be culled. Open cows at pregnancy check should be sold immediately to avoid carrying non-productive animals through the winter. The culling rate should be balanced with replacement heifer development to maintain herd size and genetic progress.

## Herd Health and Disease Prevention

Disease outbreaks can devastate profitability through mortality, reduced performance, treatment costs, and market discounts. A preventive health program is essential for maintaining productivity and minimizing losses.

### Vaccination Protocols

Producers should work with a veterinarian to develop vaccination protocols based on local disease risks. Core vaccines for beef cattle include those for clostridial diseases, bovine respiratory disease complex, and leptospirosis. The Merck Veterinary Manual provides detailed vaccination schedules and disease prevention guidelines (www.merckvetmanual.com/management-and-nutrition).

### Parasite Control

Internal and external parasites reduce feed efficiency, weight gain, and reproductive performance. A 2023 study in Animals documents the economic impact of parasitism on [beef cattle production](/knowledge/animal-farming/beef-cattle/beef-cattle-production-systems-economics-and-sustainability), including reduced weight gains, decreased milk production, and increased mortality (https://pubmed.ncbi.nlm.nih.gov/37238028). Producers should implement integrated parasite management programs that include strategic deworming, pasture rotation, and monitoring of fecal egg counts.

### Biosecurity

Biosecurity measures reduce the risk of introducing infectious diseases into the herd. New animals should be quarantined for 30 days and tested for diseases such as bovine viral diarrhea and Johne's disease before introduction. The USDA National Agricultural Library provides resources on animal health and welfare practices that support biosecurity (www.nal.usda.gov/animal-health-and-welfare).

### Genomic Selection for Disease Resistance

A 2025 study published in Animals titled "Application of Genomic Selection in Beef Cattle Disease Prevention" explores the use of genomic testing to identify animals with genetic resistance to specific diseases (https://pubmed.ncbi.nlm.nih.gov/39858277). While still emerging, this approach may allow producers to select for improved health without relying solely on vaccines and treatments.

## Forage and Grazing Management

Forage is the most cost-effective feed source for beef cattle. Proper grazing management improves forage quality and quantity, reduces feed costs, and supports animal health.

### Rotational Grazing

Rotational grazing involves moving cattle between paddocks to allow forage regrowth and prevent overgrazing. This system can increase forage utilization by 30-50% compared to continuous grazing. The USDA Natural Resources Conservation Service offers technical assistance and cost-share programs for implementing rotational grazing systems (www.nrcs.usda.gov).

### Forage Quality Testing

Forage quality varies with plant species, maturity at harvest, and storage conditions. Producers should test hay and silage for protein, energy, and mineral content to formulate balanced rations. Overfeeding supplements to compensate for poor-quality forage increases costs unnecessarily.

### Drought Management

Drought reduces forage availability and quality, forcing producers to either reduce herd size or purchase supplemental feed. Developing a drought management plan before a crisis occurs allows producers to make strategic decisions such as early weaning, culling low-performing cows, or leasing additional pasture. The Food and Agriculture Organization provides guidance on drought preparedness and risk management for livestock producers (www.fao.org/animal-production/en).

## Records and Measurement Systems

Accurate records are essential for measuring profitability and identifying areas for improvement. Without records, producers cannot calculate cost of production per head, track weaning weights, or evaluate reproductive performance.

### Essential Records

- Individual animal identification (ear tags, tattoos, or electronic ID)
- Breeding and calving dates
- Weaning weights and sale weights
- Feed purchases and forage production
- Veterinary treatments and costs
- Mortality and culling reasons

### Record-Keeping Tools

Producers can use paper records, spreadsheets, or specialized software to track production and financial data. A 2021 study published in the Joint 6th International Conference on Digital Arts, Media and Technology titled "Requirements Analysis and Design for Thai Beef-Cattle Farm Management System" highlights the need for farm management systems that integrate production records with financial data (https://doi.org/10.1109/ECTIDAMTNCON51128.2021.9425779). Producers should choose a system that is practical for their operation and use it consistently.

### Benchmarking

Comparing your operation's performance to industry benchmarks helps identify strengths and weaknesses. The Standardized Performance Analysis (SPA) program, described in a 1995 study published in The Veterinary Clinics of North America titled "Seedstock beef cattle: SPA," provides a framework for measuring and comparing production and financial performance (https://pubmed.ncbi.nlm.nih.gov/7584819). Producers can use SPA data to set realistic improvement targets.

## Common Failure Patterns and How to Avoid Them

Several recurring management failures reduce beef cattle profitability. Recognizing these patterns allows producers to take corrective action before losses accumulate.

### Failure to Track Costs

Many producers do not know their actual cost of production per head. Without this information, they cannot determine their breakeven price or make informed marketing decisions. Solution: implement a simple record-keeping system and calculate cost of production annually.

### Overstocking Pasture

Carrying more animals than the forage base can support leads to overgrazing, reduced weaning weights, and increased feed costs. Solution: calculate carrying capacity based on forage production and adjust herd size accordingly.

### Neglecting Bull Fertility

Using bulls without breeding soundness examinations reduces pregnancy rates and extends the calving season. Solution: test all bulls 30-60 days before breeding and replace subfertile animals.

### Inconsistent Health Programs

Skipping vaccinations or deworming treatments increases disease risk and reduces performance. Solution: develop a written health protocol with a veterinarian and follow it consistently.

### Poor Marketing Timing

Selling cattle when prices are low reduces revenue per head. Solution: study seasonal price patterns and use risk management tools to lock in favorable prices.

## Welfare and Safety Considerations

Animal welfare and worker safety are integral to profitable [beef cattle production](/knowledge/animal-farming/beef-cattle/beef-cattle-production-systems-economics-and-sustainability). Stressed animals perform poorly, and workplace injuries increase costs and reduce productivity.

### Animal Welfare

Cattle that are handled calmly and humanely have lower stress hormone levels, better immune function, and improved weight gain. Facilities should be designed to minimize stress during handling, loading, and transport. The USDA National Agricultural Library provides resources on animal welfare standards and best practices (www.nal.usda.gov/animal-health-and-welfare).

### Worker Safety

Beef cattle handling is inherently dangerous. Workers should be trained in safe handling techniques, proper use of equipment, and emergency procedures. Facilities should include escape routes, non-slip flooring, and adequate lighting. Producers should carry appropriate liability insurance and workers' compensation coverage.

### [Food Safety](/knowledge/bacteria/livestock-bacteria/cooking-chicken-bacteria-prevention)

Producers have a responsibility to produce beef that is safe for consumers. This includes following withdrawal periods for pharmaceuticals, maintaining clean facilities, and preventing contamination during transport. The Food and Agriculture Organization provides guidance on [food safety](/knowledge/bacteria/livestock-bacteria/cooking-chicken-bacteria-prevention) practices in livestock production (www.fao.org/animal-production/en).

## Limitations and Professional Escalation Criteria

Beef cattle profitability is influenced by factors beyond the producer's control, including weather, market prices, and input costs. Producers should recognize when they need professional assistance to address complex problems.

### When to Consult a Veterinarian

- Disease outbreaks with high morbidity or mortality
- Reproductive problems such as low pregnancy rates or high abortion rates
- Unexplained weight loss or poor performance in multiple animals
- Suspected toxic plant ingestion or chemical contamination

### When to Consult an Agricultural Economist

- Developing a long-term business plan or expansion strategy
- Evaluating the financial impact of alternative production systems
- Analyzing price risk management options
- Applying for loans or government programs

### When to Consult a Nutritionist

- Formulating rations for feedlot cattle or high-producing cows
- Diagnosing nutritional deficiencies or imbalances
- Developing supplementation programs for poor-quality forages
- Managing metabolic disorders such as milk fever or grass tetany

### When to Consult a Geneticist

- Selecting bulls or semen for specific production goals
- Interpreting genomic test results
- Developing a breeding program to improve multiple traits
- Managing inbreeding in purebred herds

## Frequently Asked Questions

### Can you make money in the cattle business?

Yes, but profitability depends on controlling costs, achieving high reproductive rates, and marketing cattle at favorable prices. Producers who track their cost of production per head and compare it to market prices can identify whether their operation is profitable. A 2022 study published in PLOS ONE titled "Evaluating profitability of [beef cattle farming](/knowledge/animal-farming/beef-cattle/beef-cattle-farming-forage-reproduction-calving-health-signals-and-herd-management) and its determinants among smallholder beef cattle farmers in the Baljovan District of Khatlon region, Tajikistan" found that access to credit, herd size, and feed costs were significant determinants of profitability (https://doi.org/10.1371/journal.pone.0274391). Profit margins in beef cattle are typically thin, and producers must manage both production and price risk.

### What is a beef cattle enterprise budget?

A beef cattle enterprise budget is a financial plan that estimates all revenues and expenses for a specific production system over a defined period, usually one year. It includes income from calf sales, cull cows, and any other products, as well as costs for feed, veterinary care, labor, and facilities. The budget calculates net return per head and per acre, allowing producers to evaluate the financial viability of different management options. Enterprise budgets are essential for making informed decisions about herd size, marketing strategies, and investment in improvements.

### How do cattle prices affect profitability?

Cattle prices directly determine revenue per head, which is the primary source of income for beef producers. When prices are high, even operations with average production efficiency can be profitable. When prices are low, only producers with low costs of production and high efficiency can maintain positive margins. Producers cannot control market prices, but they can manage price risk through forward contracts, futures hedging, and Livestock Risk Protection insurance. Understanding historical price patterns and seasonal trends helps producers time their sales to capture higher prices.

### What is the most important factor in beef cattle profitability?

Reproductive performance is widely considered the most important factor because it determines the number of calves available for sale. A cow that fails to wean a calf each year generates no revenue but incurs full annual costs. Improving pregnancy rates from 80% to 90% increases the number of calves sold by 12.5% without increasing fixed costs. Feed efficiency and weaning weight are also critical, but they cannot compensate for low reproductive rates. Producers should prioritize reproductive management as the foundation of profitability.

### How can I reduce feed costs in my beef cattle operation?

Feed costs can be reduced by maximizing the use of homegrown forages, improving pasture utilization through rotational grazing, and purchasing feed ingredients in bulk when prices are favorable. Testing forage quality allows producers to supplement only what is needed instead of overfeeding. Reducing waste through proper feeder design and management also lowers costs. In some regions, alternative feed sources such as crop residues, byproducts from food processing, or cover crops can replace more expensive traditional feeds.

### What records should I keep for beef cattle profitability?

Essential records include individual animal identification, breeding and calving dates, weaning weights, sale weights and prices, feed purchases and forage production, veterinary treatments and costs, and mortality and culling reasons. Financial records should track all income and expenses by category. Producers should calculate cost of production per head annually and compare it to industry benchmarks. Records should be maintained consistently and reviewed regularly to identify trends and areas for improvement.

### How does herd health affect profitability?

Herd health directly affects profitability through mortality, reduced performance, treatment costs, and market discounts. Disease outbreaks can cause significant losses, while chronic health problems such as parasite burdens reduce feed efficiency and weight gain. A preventive health program that includes vaccination, parasite control, and biosecurity measures reduces the risk of disease and supports optimal performance. The cost of prevention is typically much lower than the cost of treating disease outbreaks.

### What is the role of genetics in beef cattle profitability?

Genetics influence growth rate, feed efficiency, reproductive performance, carcass quality, and disease resistance. Selecting bulls and replacement heifers with superior genetics for economically important traits can improve profitability over time. Genomic testing allows producers to identify animals with high genetic potential for specific traits, including feed efficiency and disease resistance. However, genetics must be matched to the production environment and management system. High-growth genetics require adequate nutrition to express their potential, and maternal traits must be balanced with growth traits to maintain reproductive efficiency.

## Related Farming Guides

- [Beef Cattle Backgrounding Management](/knowledge/animal-farming/beef-cattle/beef-cattle-backgrounding-management)
- [Beef Cattle Forage Budgeting](/knowledge/animal-farming/beef-cattle/beef-cattle-forage-budgeting)
- [Beef Cattle Manure Management](/knowledge/animal-farming/beef-cattle/beef-cattle-manure-management)
- [Beef Cattle Marketing Records](/knowledge/animal-farming/beef-cattle/beef-cattle-marketing-records)
- [Beef Cattle Mud Management](/knowledge/animal-farming/beef-cattle/beef-cattle-mud-management)

## Related Clinical & Scientific Guides

* [Cattle Head Gate Selection and Adjustment](/knowledge/animal-farming/beef-cattle/cattle-head-gate-selection-and-adjustment)
* [Beef Cattle Handling Facility Flow](/knowledge/animal-farming/beef-cattle/beef-cattle-handling-facility-flow)
* [Beef Cattle Maternity Pen Design: Comfort and Monitoring](/knowledge/animal-farming/beef-cattle/beef-cattle-maternity-pen-design-comfort-monitoring)


## References and Further Reading

- [www.nrcs.usda.gov](https://www.nrcs.usda.gov/)
- [www.merckvetmanual.com](https://www.merckvetmanual.com/management-and-nutrition)
- [FAO Animal Production and Health](https://www.fao.org/animal-production/en). Food and Agriculture Organization of the United Nations.
- [Animal Health and Welfare](https://www.nal.usda.gov/animal-health-and-welfare). USDA National Agricultural Library.
- [The Economic Impact of Parasitism from Nematodes, Trematodes and Ticks on Beef Cattle Production.](https://pubmed.ncbi.nlm.nih.gov/37238028). Animals : an open access journal from MDPI, 2023.
- [Seedstock beef cattle: SPA.](https://pubmed.ncbi.nlm.nih.gov/7584819). The Veterinary clinics of North America. Food animal practice, 1995.
- [Telomere Dynamics in Livestock.](https://pubmed.ncbi.nlm.nih.gov/37997988). Biology, 2023.
- [Fertility management of bulls to improve beef cattle productivity.](https://pubmed.ncbi.nlm.nih.gov/27173954). Theriogenology, 2016.
- [Genetics and genomics of reproductive performance in dairy and beef cattle.](https://pubmed.ncbi.nlm.nih.gov/24703258). Animal : an international journal of animal bioscience, 2014.
- [Application of Genomic Selection in Beef Cattle Disease Prevention.](https://pubmed.ncbi.nlm.nih.gov/39858277). Animals : an open access journal from MDPI, 2025.
- [Profitable beef cattle production on rehabilitated mine lands](https://doi.org/10.1201/9781351247337). Spoil to Soil Mine Site Rehabilitation and Revegetation, 2017.
- [Requirements Analysis and Design for Thai Beef-Cattle Farm Management System](https://doi.org/10.1109/ECTIDAMTNCON51128.2021.9425779). 2021 Joint 6th International Conference on Digital Arts Media and Technology with 4th Ecti Northern Section Conference on Electrical Electronics Computer and Telecommunication Engineering Ecti Damt and Ncon 2021, 2021.
- [Competitiveness and Impact of Government Policies on Beef Cattle Business in East Kolaka District, Indonesia: A Policy Analysis Matrix Approach](https://doi.org/10.21059/buletinpeternak.v49i2.105016). Buletin Peternakan, 2025.
- [Management of beef cattle production in Malaysia: A step forward to sustainability](https://doi.org/10.3844/ajassp.2016.976.983). American Journal of Applied Sciences, 2016.
- [Evaluating profitability of beef cattle farming and its determinants among smallholder beef cattle farmers in the Baljovan District of Khatlon region, Tajikistan](https://doi.org/10.1371/journal.pone.0274391). Plos One, 2022.

> This article is educational and is not a substitute for veterinary diagnosis, treatment, public-health guidance, or regulatory reporting.


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