Why Insects Fly Toward Lights: The Science of Phototaxis
The sight of insects swarming around a porch light or streetlamp is familiar to anyone who spends time outdoors at night. This behavior, called positive phototaxis, is the directed movement of an organism toward a light source. While the phenomenon is well documented, the underlying mechanisms are more complex than a simple attraction to brightness. Current research points to several interacting explanations, including navigation disruption, sensory confusion, and species-specific visual adaptations. Understanding why insects fly toward lights matters for pest management, conservation of nocturnal pollinators, and design of lighting systems that minimize ecological harm.
This article examines the leading scientific hypotheses for insect attraction to artificial light, compares how different insect groups respond, and explains practical implications for lighting decisions and insect monitoring. The focus is on what researchers have measured and observed, with attention to where evidence remains incomplete.
The Leading Hypotheses for Flight-to-Light Behavior
Navigation Confusion and the Dorsal Light Response
The most widely cited explanation for why insects fly toward artificial lights involves disruption of their natural navigation systems. Many nocturnal insects maintain straight flight paths by keeping a bright visual cue, such as the moon or a distant celestial light source, at a constant angle relative to their eyes. This is sometimes called the transverse orientation mechanism. When an insect encounters a nearby artificial light, the angle to that light changes rapidly as the insect moves, unlike the effectively parallel rays from the moon. The insect continuously adjusts its heading to keep the light at the same visual angle, resulting in a spiral path that draws it ever closer to the source.
A related mechanism involves the dorsal light response. Many insects maintain their orientation in flight by keeping the brightest part of their visual field above them. A bright artificial light on the ground or at eye level can override the natural skyward brightness gradient, causing the insect to tilt and steer toward the light. This response is a disruption of the sensory feedback the insect uses to stay upright and on course.
Evidence for navigation disruption comes from studies of moths in urban environments. Research on spindle ermine moths found that urban populations have evolved a reduced flight-to-light response compared to rural populations, and this change correlates with smaller wings. The authors of that study support the reduced mobility hypothesis, which states that reduced mobility in urban areas is associated with specific morphological changes in the flight apparatus. This finding suggests that flight-to-light behavior carries substantial fitness costs, strong enough to drive evolutionary change in urban moth populations. The study was published in Biology Letters in 2024 and is available through PubMed.
Positive Phototaxis as a Sensory Response
Positive phototaxis is the direct movement toward a light source, and it is not limited to nocturnal insects. Many insects show positive phototaxis to specific wavelengths of light, and this response is exploited in insect monitoring and control. The response is not uniform across species or even within a species across different light conditions.
Research on the whitefly Bemisia tabaci and its predator Serangium japonicum demonstrated that phototaxis depends on wavelength, light intensity, and temperature. In Y-tube assays, B. tabaci consistently showed strong positive phototaxis. At irradiances from 100 to 800 µW cm⁻², positive phototaxis was highest under 480 nm light, while at 1000 µW cm⁻² it was highest under 520 nm light. The predator S. japonicum showed negative phototaxis under several conditions, with avoidance strongest under 440 nm at 200 µW cm⁻² and under 400 nm at higher irradiances. This avoidance weakened as temperature increased. The study, published in Insects in 2026, is available at DOI 10.3390/insects17070661.
This research has direct practical implications. LED-based trapping for pest management must account for the fact that light conditions attracting pests may also repel or attract beneficial predators. The authors concluded that selective LED trapping should be deployed as a temperature-aware strategy instead of as a single fixed setting.
Escape Behavior and the Confusion of Bright Surfaces
Some insects may fly toward light because bright surfaces resemble open space or escape routes. Insects trapped in dark enclosures often move toward the brightest available area, a behavior that may help them find exits in natural settings. Artificial lights at night may be perceived as gaps in vegetation or openings in the canopy, drawing insects toward them.
This explanation overlaps with the sensory confusion hypothesis. Insects have evolved visual systems optimized for their natural environments, and artificial lights present stimuli that do not occur in nature. The insect responds using rules that work in natural settings but produce maladaptive outcomes around human-made lights.
Blue-Green Opponency and Host Detection
Research on biting flies offers insight into how specific visual channels drive attraction to certain colors. Diurnal biting flies are strongly attracted to blue objects, a behavior widely exploited for fly control. A 2023 study in Proceedings of the Royal Society B tested three hypotheses for this attraction: blue objects resemble animal hosts, blue surfaces resemble shaded resting places, and blue attraction is a by-product of attraction to polarized light. The researchers computed fly photoreceptor signals from leaf and animal integument reflectance spectra and trained artificial neural networks to distinguish animals from leaf backgrounds and shaded from unshaded surfaces. The trained networks discriminated animals from leaves based on blue-green photoreceptor opponency and commonly misclassified blue objects as animals. The authors concluded that blue-green opponency is the most effective means of discriminating animals from leaf backgrounds using a fly's sensory information, and that blue objects resemble animal hosts through such mechanisms. The study is available through PubMed.
This finding illustrates a broader principle: insect attraction to light and color is mediated by the same sensory channels used for essential behaviors like host finding. Artificial lights that stimulate these channels can trigger responses that are adaptive in natural contexts but maladaptive around human structures.
At a Glance: Insect Group Responses to Artificial Light
| Insect Group | Typical Response to Artificial Light | Likely Mechanism | Evidence Source |
|---|---|---|---|
| Moths (Lepidoptera) | Strong positive phototaxis, often circling lights | Navigation disruption, dorsal light response | PubMed: Evolutionary change in flight-to-light response in urban moths |
| Glow-worms (Lampyris noctiluca) | Males avoid white light, reduced mate finding | Light aversion, disruption of bioluminescent signaling | PubMed: Artificial light impairs local attraction to females in male glow-worms |
| Whiteflies (Bemisia tabaci) | Strong positive phototaxis, wavelength dependent | Spectral sensitivity, peak response at 480 nm | DOI 10.3390/insects17070661 |
| Biting flies (diurnal) | Attracted to blue objects | Blue-green opponency resembling animal hosts | PubMed: Why are biting flies attracted to blue objects |
| Midges (Diptera) | Positive phototaxis, earlier activity than moths | Taxon-specific timing, spectrum dependent | Europe PMC: Timing of attraction to light of nocturnal insects |
| Locusts (Locusta migratoria manilensis) | Aggregation under combined spectral and polarized light | Polarotaxis combined with phototaxis | DOI 10.3390/insects17060569 |
Spectrum and Timing: How Light Color Changes Insect Behavior
Wavelength-Dependent Phototaxis
Insects do not respond equally to all wavelengths of light. Their compound eyes contain photoreceptors with peak sensitivities in different spectral regions, and these sensitivities vary among species. Understanding which wavelengths attract which insects is essential for designing lighting that minimizes ecological disruption while still serving human needs.
The study on Bemisia tabaci and Serangium japonicum provides a clear example of wavelength-dependent responses. Whiteflies showed peak positive phototaxis at 480 nm (blue) at most irradiance levels, shifting to 520 nm (green) at the highest irradiance tested. The predatory ladybird S. japonicum showed strongest avoidance at 400 nm (violet) and 440 nm (blue) depending on irradiance. This differential response creates an opportunity for selective trapping: the authors identified 400 nm at 600 µW cm⁻² as the best predator-safe setting at 20 to 30 °C, and 440 nm at 600 µW cm⁻² at 35 °C. These findings are reported in Insects 2026.
Temporal Patterns in Phototaxis
The timing of insect attraction to light varies by taxon and by light spectrum. A 2026 study in the Netherlands used camera-light traps with LED strips emitting four monochromatic spectra and one dichromatic spectrum to assess phototactic activity over twelve nights at four forested locations. Phototactic activity intensified from early to middle night but varied with taxon. Diptera (midges) exhibited an earlier center of phototactic activity than Lepidoptera (moths). Midges were attracted earlier to ultraviolet and red light and later to green and amber light. The timing of moth phototaxis was unaffected by light color. The study is available through Europe PMC.
These findings support the potential for time-scheduled, spectrally adjustable lighting systems. Reducing light emission in the early and middle parts of the night, when phototactic activity peaks, could mitigate negative effects on nocturnal insects.
Developmental Light Environment and Adult Behavior
The light environment experienced during development can shape adult phototaxis. A 2026 study on Drosophila melanogaster reared flies under white, blue, green, yellow, or red light from egg to eclosion. Flies developed under color-filtered lights exhibited wavelength-congruent phototactic preferences, spending more time near the wavelength corresponding to their developmental light environment. Flies developed under red light showed significantly higher movement and active speeds than those developed under short-wavelength or white light. After five days of recovery under white light, both wavelength-specific phototactic preferences and enhanced locomotion were largely diminished, indicating that these effects are plastic and reversible. The study is available at DOI 10.1016/j.beproc.2026.105383.
This plasticity has implications for laboratory rearing and for understanding how insect populations adapt to artificially lit environments. Insects that develop in lit conditions may show different behavioral responses than those that develop in darkness, complicating predictions about light trap effectiveness.
Artificial Light at Night and Insect Physiology
Disruption of Diapause
Artificial light at night does more than attract insects. It can disrupt fundamental physiological processes, including diapause, the dormant state many insects enter to survive winter. Many insects regulate diapause in response to photoperiod, with short day length signaling the approach of winter and inducing diapause. Artificial light at night can disrupt these photoperiodic cues and inhibit diapause induction.
A 2025 review in the Journal of Insect Physiology examined this disruption, summarizing recent empirical studies on ALAN-induced diapause inhibition and outlining the physiological and molecular genetic mechanisms involved, which remain only partially understood. The review also discussed potential factors limiting adaptation to ALAN-induced diapause disruption and presented perspectives from physiological, evolutionary, and applied entomological standpoints. It emphasized the need to integrate existing knowledge of diapause regulation with field and laboratory studies. The review is available through PubMed.
Case Study: Aedes albopictus
A 2024 study in The Science of the Total Environment examined the effect of artificial light at night on diapause in Aedes albopictus, an important disease vector. The researchers studied diapause characteristics of different geographical strains under ALAN interference and explored molecular mechanisms through RNA-seq. Aedes albopictus of the same geographic strain showed lower diapause incidence when exposed to ALAN. In the short photoperiod induced diapause of temperate strain Beijing and subtropical strain Guangzhou, ALAN disturbance reduced the egg diapause rate and increased the egg hatching rate. ALAN disturbance also shortened the life cycle of Aedes albopictus eggs after hatching. The study is available through PubMed.
This research has public health implications. If ALAN reduces diapause and increases hatching rates, it could extend the active season of disease vectors in urban areas, potentially increasing transmission risk. The authors noted that understanding ALAN effects on diapause could provide a reference for prevention and control of infectious diseases mediated by Aedes albopictus.
Disruption of Bioluminescent Communication
Glow-worms rely on bioluminescence for reproduction. Females emit light to attract males, and artificial light at night can interfere with this signal. A 2023 study in The Journal of Experimental Biology quantified the effect of white illumination on male glow-worms' ability to reach a female-mimicking LED within a Y-maze. As illumination intensity increased, the proportion of males reaching the LED declined. Brighter illumination also increased the time taken to reach the LED, because males spent more time in the central arm of the maze and with their head retracted beneath their head shield. These effects reversed rapidly when illumination was removed, suggesting that male glow-worms are averse to white light. The study is available through PubMed.
An earlier 2020 study in the same journal compared trap efficacy in the presence or absence of a white artificial light source. Illuminated traps attracted fewer males than traps in the dark. Illuminated traps closer to the light source attracted fewer males than those further away, while traps in the dark attracted similar numbers of males up to 40 m from the light source. The authors concluded that ALAN impedes females' ability to attract males, with the effect increasing with light intensity, potentially affecting fecundity and long-term population survival. The study is available through PubMed.
A related study on white LED light intensity and color temperature in glow-worms found that intensity, but not color temperature, interferes with mate-finding by Lampyris noctiluca males. The bibliographic record is available at DOI 10.1007/s10841-021-00304-z.
Nocturnal Pollination and Light Pollution
The Special Niche of Night
The night is a distinct ecological niche characterized by dim light, lower temperatures, and higher humidity compared to the day. Several animal groups have transitioned from day to night activity and acquired unique adaptations for nocturnal function. Many plant species bloom at night, possibly as a response to aridity to prevent excessive water loss through evapotranspiration or to protect pollen from heat stress. Nocturnal pollinators have visual adaptations for dim light conditions but may trade off vision against olfaction when dependent on nectar-rewarding and scented flowers. Some nocturnal pollinators use carbon dioxide and humidity cues from freshly opened flowers as indicators of nectar-rich resources. Some endothermic nocturnal insect pollinators are attracted to thermogenic flowers within which they remain to obtain heat as a reward. A 2018 review in The Yale Journal of Biology and Medicine covers these mechanisms and notes that excessive and inappropriate illumination from anthropogenic activities has resulted in significant light pollution that undermines life processes governed by dim light. The review is available through PubMed.
Positive Effects in a Specialized Mutualism
Not all effects of artificial light on plant-pollinator systems are negative. A 2021 study in Integrative and Comparative Biology examined the obligate mutualism between yucca moths (Tegeticula maculata maculata) and yucca plants (Hesperoyucca whipplei). The researchers deployed field-placed light towers during peak yucca flowering and compared moth captures between dark-controlled and light-treated trials. Adult moth abundance was much higher when light was present, suggesting that ALAN may alter this diurnal moth's activity patterns to expand their temporal niche into the night. Both larva and fruit recruitment increased with skyglow, and fruit set also increased with direct lighting, though the relationship was weaker. Larva recruitment did not change when exposed to a gradient of direct light, which may reflect effects of ALAN on moth physiology, such as disrupted female oviposition. The study is available through PubMed.
This study demonstrates that ALAN effects are context dependent. In this tightly co-evolved mutualism, ALAN positively influenced the fitness of both plants and moths, but the benefits to each species may not be equal. The authors noted that larva recruitment did not increase with direct light despite higher adult abundance, suggesting that light may disrupt oviposition behavior even when it attracts adults.
Visual Adaptations and Seasonal Variation
Compound Eye Morphology
Insect visual systems are adapted to their ecological niches, and these adaptations can influence phototaxis. A 2026 study examined the compound eye morphology of Scythris sinensis, a diurnal moth with distinct spring and autumn forms. Using scanning electron microscopy, the researchers found that the eyes are ellipsoidal and symmetrical, consisting of hexagonal ommatidia with convoluted folds but lacking corneal nipples. Female moths had slightly larger compound eyes than males. Males emerging in autumn typically possessed a higher number of ommatidia (589 to 675) than males emerging in spring (492 to 698). This finding suggests an adaptation specific to the autumn period, potentially enhancing visual search efficiency in response to seasonal light variations and a constrained reproductive schedule. The study is available at DOI 10.3390/insects17070702.
Seasonal variation in eye morphology means that the same species may show different phototactic responses at different times of year. Monitoring programs that use light traps should account for this variation when interpreting catch data.
Polarized Light and Aggregation
Some insects respond to light polarization in addition to intensity and wavelength. A 2026 study on Locusta migratoria manilensis, a locust species that causes frequent outbreaks in China, examined the synergistic effects of spectral light and linear polarized light on visual aggregation. The results showed that the coupling of unpolarized and polarized components within partially polarized light determines the effectiveness of polarized spectral vector illumination on locust aggregation sensitivity. Among tested conditions, unpolarized violet light combined with linearly polarized orange light at a 270 degree vector angle produced the strongest time-dependent effect, followed by unpolarized orange light combined with linearly polarized violet light at 0 degrees. The combined stimulation of spectral and polarized light triggered aggregation through polarization-enhanced polarotaxis and spectrum-driven phototaxis. The study is available at DOI 10.3390/insects17060569.
These findings provide a basis for improved optical control strategies for locusts, using partially polarized light to trap locusts, promote aggregation, and interfere with navigation.
Practical Applications for Lighting and Insect Management
Designing Light Traps
Light traps are widely used for insect monitoring and control. The effectiveness of a trap depends on matching the light spectrum to the target species while minimizing impacts on nontarget organisms. The research on Bemisia tabaci and Serangium japonicum provides a model for this approach. By testing multiple wavelength-irradiance combinations, the researchers identified settings that attracted the pest while repelling or not affecting the predator. The temperature dependence of these responses means that trap settings may need adjustment seasonally. The study is available at DOI 10.3390/insects17070661.
Reducing Unwanted Insect Attraction
For homeowners and facility managers, reducing insect attraction to lights involves several practical steps. Choosing warm-colored lights with lower ultraviolet output can reduce attraction for many insect groups, though the effectiveness varies by taxon. The temporal study from the Netherlands showed that midges were attracted earlier to ultraviolet and red light and later to green and amber light, while moth attraction timing was unaffected by color. This means that no single lighting choice will eliminate attraction for all insects. The study is available through Europe PMC.
Light curfews, or turning off lights during peak insect activity periods, may be more effective than spectral tuning alone. The same study found that phototactic activity intensified from early to middle night, suggesting that reducing emissions during this window could have disproportionate benefits.
Monitoring and Assessment
For researchers and pest management professionals, assessing the impact of artificial light on local insect populations requires systematic observation. Key measurements include:
- Species composition and abundance at light sources over time
- Timing of peak activity relative to lighting schedules
- Behavioral observations, such as circling versus direct approach
- Reproductive outcomes, such as mating success or oviposition rates
- Physiological indicators, such as diapause incidence in vector species
The glow-worm studies demonstrate that field observations alone may miss behavioral impacts. The 2023 Y-maze study showed that ALAN prevents male glow-worms from reaching females and increases the time they take to reach females and the time they spend avoiding exposure to light. The authors noted that this raises the possibility that ALAN has similar behavioral impacts on other insect species that remain undetected in field experiments. The study is available through PubMed.
Common Failure Patterns in Interpreting Flight-to-Light Behavior
Assuming Uniform Attraction
A common error is assuming that all insects are equally attracted to all lights. The evidence shows substantial variation by taxon, wavelength, intensity, temperature, and time of night. The whitefly study demonstrated that even within a species, the preferred wavelength shifts with irradiance. The temporal study showed that different taxa have different activity peaks. Generalizations about insect attraction to light should be treated with caution.
Ignoring Context-Dependent Effects
The yucca moth study showed that ALAN can have positive effects on some species in some contexts. Assuming that all artificial light is harmful to all insects is not supported by the evidence. The effects depend on the species, the light characteristics, and the ecological context.
Overlooking Behavioral Plasticity
The Drosophila study demonstrated that phototactic preferences can be shaped by developmental light environment and can reverse after recovery under different conditions. This plasticity means that laboratory results may not predict field behavior, and that populations may adapt to lit environments over time. The urban moth study provides evidence for evolutionary change in flight-to-light response, with urban populations showing reduced attraction and associated morphological changes. Both studies are available through PubMed and DOI 10.1016/j.beproc.2026.105383.
Limitations of Current Knowledge
Incomplete Mechanistic Understanding
While several hypotheses explain flight-to-light behavior, the precise mechanisms remain unclear for many species. The 2025 diapause review noted that the physiological and molecular genetic mechanisms involved in ALAN-induced diapause disruption remain only partially understood. The authors called for integrating existing knowledge of diapause regulation with field and laboratory studies. The review is available through PubMed.
Gaps in Field Detection
The 2023 glow-worm study highlighted a critical limitation: behavioral impacts of ALAN may remain undetected in field experiments. The Y-maze study revealed that male glow-worms spend more time avoiding light exposure, a behavior that would be difficult to observe in natural settings. This suggests that the true impact of ALAN on insect populations may be underestimated. The study is available through PubMed.
Limited Taxonomic Coverage
Research on flight-to-light behavior has focused on a relatively small number of species, including moths, glow-worms, whiteflies, fruit flies, and a few others. The extent to which findings generalize across the enormous diversity of insects is unknown. The temporal study from the Netherlands examined multiple taxa but was limited to forested locations in a single country. The study is available through Europe PMC.
Safety and Regulatory Context
Public Health Considerations
The Aedes albopictus study has direct public health relevance. If artificial light at night reduces diapause and increases egg hatching rates, it could extend the active season of this disease vector in urban areas. The authors noted that understanding ALAN effects on diapause could provide a basis for new vector control strategies and a reference for prevention and control of infectious diseases mediated by this species. The study is available through PubMed.
Occupational and Operational Safety
For professionals working with light traps or managing lighting systems, standard electrical safety practices apply. Light traps should be installed according to manufacturer specifications, with proper grounding and protection from moisture. When working at night, use additional lighting for safe navigation, and be aware that insect activity may attract predators or create slippery surfaces near traps.
Escalation Criteria
Professionals should consider escalating to specialized expertise when:
- Light trap catches show unexpected species composition changes that may indicate broader ecological impacts
- Vector species show extended activity seasons that may require public health consultation
- Pest populations do not respond to light-based control measures as expected, suggesting resistance or behavioral adaptation
- Lighting modifications intended to reduce insect attraction do not produce measurable changes in insect activity
Frequently Asked Questions
What insects fly toward lights?
Many nocturnal insects fly toward artificial lights, including moths, midges, beetles, and true flies. The response varies by species and light characteristics. Moths are among the most conspicuous, often circling lights persistently. Midges show positive phototaxis with timing that varies by light spectrum. Some diurnal insects, such as yucca moths, may also be attracted to artificial light at night, expanding their activity into the nocturnal period. Evidence for these patterns comes from studies on urban moths, midges, and yucca moths, available through PubMed, Europe PMC, and PubMed.
What insects fly and sting?
The insects most commonly associated with both flight and stinging are in the order Hymenoptera, which includes bees, wasps, and ants. These insects are primarily diurnal and are not typically attracted to artificial lights at night, though some species may be drawn to lights under certain conditions. The evidence reviewed here does not address stinging insects directly. For information on stinging insect behavior, consult local extension services or pest management professionals.
Why do moths circle lights?
Moths circling lights is consistent with navigation disruption. The insect attempts to maintain a constant visual angle to the light source, which produces a spiral flight path around the light. This behavior disrupts their general activity focused on finding resources such as mating partners and increases predation risk. Urban moth populations have evolved reduced flight-to-light responses, supporting the idea that this behavior carries substantial fitness costs. The evidence is reported in Biology Letters 2024.
Are all insects attracted to light?
No. Some insects show negative phototaxis, moving away from light. The predatory ladybird Serangium japonicum showed negative phototaxis under several wavelength and irradiance conditions in laboratory assays. Male glow-worms avoid white light, retracting their heads beneath their head shields when illuminated. The response to light depends on species, wavelength, intensity, temperature, and ecological context. Evidence is available at DOI 10.3390/insects17070661 and PubMed.
Does light color matter for insect attraction?
Yes. Different insect species have different spectral sensitivities, and the same species may respond differently to different wavelengths. Whiteflies showed peak attraction at 480 nm at most irradiance levels, shifting to 520 nm at the highest irradiance. Midges were attracted earlier to ultraviolet and red light and later to green and amber light. Moth attraction timing was unaffected by light color. These findings support the potential for spectrally tuned lighting to reduce unwanted insect attraction. Evidence is available at DOI 10.3390/insects17070661 and Europe PMC.
Can insects adapt to artificial light?
Yes. Urban populations of spindle ermine moths have evolved reduced flight-to-light responses compared to rural populations, and this change correlates with smaller wings. The reduced mobility hypothesis states that reduced mobility in urban areas is associated with specific morphological changes in the flight apparatus. This demonstrates that flight-to-light behavior can change through evolution when it imposes fitness costs. The evidence is reported in Biology Letters 2024.
How does artificial light affect insect reproduction?
Artificial light can disrupt reproduction in several ways. In glow-worms, white light interferes with the bioluminescent signal females use to attract males, reducing mate-finding success. The effect increases with light intensity and can be detected up to 40 m from the light source. In Aedes albopictus, artificial light at night reduces diapause incidence and increases egg hatching rates, potentially extending the active season. Evidence is available at PubMed, PubMed, and PubMed.
Can artificial light ever benefit insects?
In some contexts, yes. The yucca moth study found that adult moth abundance was much higher when light was present, and both larva and fruit recruitment increased with skyglow in this obligate mutualism. The authors suggested that ALAN may alter this diurnal moth's activity patterns to expand their temporal niche into the night. However, larva recruitment did not change with direct light exposure, which may reflect disrupted oviposition behavior. The study is available through [PubMed
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References and Further Reading
- NCBI Literature Resources. National Center for Biotechnology Information.
- PubMed. National Library of Medicine.
- Artificial light impairs local attraction to females in male glow-worms.. The Journal of experimental biology, 2023.
- The age of bright nights: Photoperiodic disruption of insect diapause by artificial light at night.. Journal of insect physiology, 2025.
- Why are biting flies attracted to blue objects?. Proceedings. Biological sciences, 2023.
- Artificial lighting impairs mate attraction in a nocturnal capital breeder.. The Journal of experimental biology, 2020.
- Dark Matters: Challenges of Nocturnal Communication Between Plants and Animals in Delivery of Pollination Services.. The Yale journal of biology and medicine, 2018.
- The effect of artificial light at night (ALAN) on the characteristics of diapause of Aedes albopictus.. The Science of the total environment, 2024.
- Evolutionary change in flight-to-light response in urban moths comes with changes in wing morphology.. Biology letters, 2024.
- Direct and Ambient Light Pollution Alters Recruitment for a Diurnal Plant-Pollinator System.. Integrative and comparative biology, 2021.
- Spectral- and Temperature-Dependent Phototaxis of <,i>,Bemisia tabaci<,/i>, and Its Predator <,i>,Serangium japonicum<,/i>,: Implications for Predator-Aware Selective LED-Based Trapping.. 2026.
- Timing of attraction to light of nocturnal insects is spectrum and taxon dependent: implications for mitigating light pollution. 2026.
- Visual Aggregation Sensitivity of the <,i>,Locusta migratoria manilensis<,/i>, Under Partially Polarized Light in Greenhouses.. 2026.
- Seasonal Dimorphism in the Compound Eye Morphology of <,i>,Scythris sinensis<,/i>, (Felder &, Rogenhofer, 1875) (Lepidoptera: Scythrididae).. 2026.
- Developmental light colour influences adult phototaxis and locomotion in Drosophila melanogaster.. 2026.
- WHY FLY NOW ? PUPA BANKS , APOSEMATISM , AND OTHER FACTORS THAT MAY EXPLAIN OBSERVED MOTH FLIGHT ACTIVITY. 2016.
- White LED light intensity, but not colour temperature, interferes with mate-finding by glow-worm (Lampyris noctiluca L.) males. Journal of Insect Conservation, 2021.
This article is educational and does not replace institutional policy, professional advice, or applicable safety and regulatory requirements.